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YOUR
PHOTOGRAPHS SUBMITTED FOR T-LIST DISCUSSIONS:
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T-List
#: TL07-2006-04-18
Subject: T. flexipes Clumping
Observations: Offset propagation.
Submitter: Harold
Holmes
Discussion: 2005
decapitation of 5 rhizomes quarter-cut in 13 sections yielded 37
offsets when observed in April 2006 (11 months).
Photograph: Harold Holmes(click on
image for medium size) LARGE
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18B. T. flexipes
18 April 2006
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18A. T. flexipes
18 April 2006
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John Gyer Writes:
I wanted to do more photo work before I started sending out micrographs,
but these have bearing on the current discussion of offset formation.
I do not have time to annotate so I'll give descriptions quickly.
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T-List
#: TL07-2006-04-21A
Subject:
Offset propagation: T. grandiflorum rhizome with
root initial, Iodine stained 2005 JFGYER
Submitter: John
Gyer
Discussion:This shows the differentiated
buds of trillium. The section is oblique through the rhizome apex
of a 3 year seedling. Two root traces are present; one at about
2o'clock and the other at about 5o'clock. THIS YEAR's STEM produced
the vascular traces at about 7o'clock. At about 10o'clock is the
differentiated bud for NEXT year's growth covered by a cataphyll.
Above that at about 11o'clock is the less differentiated bud for
growth the YEAR AFTER NEXT. Variations on this timing are possible
depending on the quality of the season and the vigor of growth.
These variations result in plants with two or more apical stems.
Differentiated buds such as these have NOT been seen behind the
rhizome apex.
Photograph: John Gyer(click on image
for medium size) LARGE
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T-List
#: TL07-2006-04-21B
Subject: Offset propagation:
T.grandiflorum Iodine Stained bud section 2005 JFGYER
Submitter: John
Gyer
Discussion: This
is a rhizome section a short distance behind the rhizome apex. Stored
starch is stained black by the Iodine. The roughly circular structure
of white filaments (no starch present) is the STELE. The vasculature
for both roots and shoots arise from the surface of this structure.
In this micrograph a root initial is seen sprouting from a section
of the stele and growing its way through the cortex toward the surface
of the rhizome. The bluish-purple color of the cells in and around
the stele at this section level is due to the metabolic state of
the cells. Since no photosynthesis was present when the sectioning
was done, the color probably represents cells that are converting
stored starch into the energy needed for growth. Since: a. no differentiated
buds were seen on the rhizome surface; b. normal roots (and shoots)
form from cambial tissue associated with the stele; then it is highly
probable that the offsets seen in Harold's experiments also originate
in this structure from its associated cambium tissue.
Photograph: John Gyer(click on image
for medium size) LARGE
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John Gyer Clarksboro NJ USA
Trillium
Books and Publications:
- Trilliaceae resources: (print resources compiled from Susan Farmer
list)
- Samejima and Samejima. 1987. Trillium genus illustrated.
Long out of print, but worth the effort from InterLibrary Loan.
- Freeman, J.D. 1975. Revision of Trillium subgenus Phyllantherum
(Liliaceae). Brittonia 27:1-62.
The paper on sessile-flowered trilliums.
- Case and Case. 1997. Trilliums.
Considered by many to be the standard. Technical and practical.
- Jacobs and Jacobs. 1997. Trilliums in woodland and garden: American
treasures.
More oriented for the gardener. Available from the author.
- Li. 1998. The genus Paris.
In chinese.
- Henderson. 2000. Trilliums for your garden.
Very nice. Lots of pictures. Available from the author.
Links
to Trillium Articles: This will link
you to 30+ trillium articles on the Internet.
Links
to Trillium Websites:
This will link you to trillium websites on the Internet.
Your
Feedback and Submittals:
Please email me your comments ...<comments@trilliumresearch.org>.
Please email any 'trillium articles,
references, or formal or informal publications or processes' submittals
to ... <harold@trilliumresearch.org>
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